Publications from the Whitehead lab. (1977-present)
Select year of
publication:
In Press 2024 2023
2022 2021 2020 2019 2018 2017 2016 2015 2014 2013 2012 2011 2010 2009 2008 2007 2006 2005 2004 2003 2002 2001 2000 1999 1998 1997 1996 1995 1994 1993 1977-1992
Brakes, P. and H. Whitehead. In
press. Culture and conservation. In: Evolution of cetacean societies.
Edited by D. Croft, A Foote, E. Garland and S. King. University of Chicago
Press, Chicago.
Eguiguren, A., I. Avila, S. Mesnick, M. Cantor, T. Hersh, H.
Pérez-Puig, P. Rosero, L. Rendell, H. Whitehead, C. Rojas and J. Alava. In
press. Integrating cultural dimensions in sperm whale (Physeter macrocephalus)
conservation: threats, challenges, and solutions. Philosophical Transactions
of the Royal Society B
Publications 2025
Jones, L., J.M. Allen, K.C.
Balcomb, C.J. Basran, S.D. Berrow, L. Betancourt, L. Bouveret,
T.K. Boye, F. Broms, V. Chosson, P.J. Clapham, T.W.
Fernald, J.G. Frediani, B. Jann, R. Kempen, J. Lien, P. López Suárez, E.E.
Magnúsdóttir, D.K. Mattila, M.M. MacKay, N. Øien, M.H. Rasmussen, M. Rinaldi,
J. Robbins, R. Sears, R.E. Seton, O. Shamir Reynoso, M. Simon, P.T. Stevick,
G.A. Víkingsson, M. Weinrich, F.W. Wenzel, H. Whitehead, D. Young and S.K.
Todd. 2025. Ocean-basin-wide movement patterns of North Atlantic humpback
whales, Megaptera novaeangliae. Journal of Cetacean Research and Management
26. https://journal.iwc.int/index.php/jcrm/article/view/951 PDF
Rendell, L, and H. Whitehead. 2025.
The active space of sperm whale codas allows for communication within and
between social units. Journal of Experimental Biology 227 (15):
jeb247784 Journal
article
Whitehead, H., and S.F. Walmsley. 2025.
Societies of the open ocean without territories. Behavioral and Brain
Sciences 48: e74. Journal article
Feyrer, L.J., J.E. Stanistreet, C. Gomez, M. Adams,
J.W. Lawson, S.H. Ferguson, S.G. Heaslip, K.J. Lefort, E. Davidson, N.E.
Hussey, H. Whitehead and H. Moors-Murphy. 2024. Identifying important habitat
for northern bottlenose and Sowerby’s beaked whales in the western North
Atlantic. Aquatic Conservation: Marine and Freshwater Ecosystems 34:
e4064. Journal article
Hersh, T.A., A Ravignani and H. Whitehead. 2024. Cetaceans are the next frontier for vocal rhythm research. Proceedings of the National Academy of Sciences (USA) 25: e2313093121. Journal article
Tyack, P.L., C.W. Clark, P.T. Madsen, C. Walcott, H.
Whitehead, and B. Wuersig. 2024. Roger Payne
1935-2023. Marine Mammal Science e13172.
Vachon, F., L. Rendell, S. Gero and H. Whitehead.
2024. Abundance estimate of Eastern Caribbean sperm whales using large scale
regional surveys. Marine Mammal Science 40: e13116 PDF
Whitehead, H. 2024. Sperm whale clans and human
societies. Royal Society Open Science 11: 231353 Journal article
Kelly, N.E., L. Feyrer, H. Gavel, O. Trela, W. Ledwell, H. Breeze, E.C. Marotte, L. McConney and H. Whitehead. 2023. Long term trends in floating plastic pollution within a marine protected area identifies threats for endangered northern bottlenose whales. Environmental Research 227: 115686. PDF
Whitehead, H., J.K.B. Ford and A.G. Horn. 2023.
Using culturally transmitted behavior to help delineate conservation units for
species at risk. Biological Conservation 285: 110239. PDF
Zwamborn, E., N.
Boon and H. Whitehead. 2023. Collective
decision-making in aquatic mammals. Mammal Review 53: 238-253. PDF
Zwamborn, E., S.
Walmsley and H. Whitehead. 2023. Flanking female guides: collective
decision-making in long-finned pilot whales. Animal Behaviour
205: 149-159. PDF
Hersh, T.A., S. Gero, L. Rendell, M. Cantor, L. Weilgart,
M. Amano, S.M. Dawson, E. Slooten, C.M. Johnson, I. Kerr, R. Payne, A. Rogan,
R. Antunes, O. Andrews, E.L. Ferguson, C.A. Hom-Weaver, T.F. Norris, Y.M.
Barkley, K.P. Merkens, E.M. Oleson, T. Doniol-Valcroze,
J.F. Pilkington., J. Gordon, M. Fernandes, M. Guerra, L. Hickmott and H.
Whitehead. 2022. Symbolic marking in non-human cultures: evidence from sperm
whale clans. Proceedings of the National Academy of Sciences (USA)
119: e2201692119. PDF
Kowarski, K., S.
Cerchio, H. Whitehead, D. Cholewiak and H.
Moors-Murphy. 2022. Seasonal song ontogeny in western North Atlantic humpback
whales: drawing parallels with songbirds. Bioacoustics 31:450-469. Journal
article
Tavares, S.B., H. Whitehead and T. Doniol-Valcroze. 2022. Assessing social structure: a data-driven approach to define associations between individuals. Mammalian Biology 102, 551-566. Journal article
Vachon, F., A. Eguiguren,
L. Rendell, S. Gero and H. Whitehead. 2022. Distinctive, fine-scale
distribution of Eastern Caribbean sperm whale vocal clans reflects island
fidelity rather than environmental variable. Ecology and Evolution 12:e
9449. Journal
article
Vachon, F., T.A. Hersh, L. Rendell, S. Gero and H. Whitehead. 2022. Ocean nomads or island specialists? Culturally driven habitat partitioning contrasts in scale between geographically isolated sperm whale populations. Royal Society Open Science 9: 211737. http://doi.org/10.1098/rsos.211737 Journal article Video abstract
Whitehead, H., and T.A. Hersh. 2022. Posterior probabilities of membership of repertoires in acoustic clades. PLoS ONE 17(4): e0267501. https://doi.org/10.1371/journal.pone.0267501 Journal article
Whitehead, H., and M. Shin. 2022. Current global population size, post-whaling trend and historical trajectory of sperm whales. Scientific Reports 12: 19468. Journal article
Publications
2021
Brakes, P., E.L. Carroll, S.R.X.
Dall, S.A. Keith, P.K. McGregor, S.L. Mesnick, M.J. Noad, L. Rendell, M.M.
Robbins, C. Rutz, A. Thornton, A. Whiten, M.J. Whiting, L.M. Aplin, S. Bearhop, P. Ciucci, V. Fishlock, J.K.B. Ford, G. Notarbartolo di Sciara, M.P. Simmonds, F. Spina, P.R. Wade,
H. Whitehead, J. Williams and E.C. Garland. 2021. A deepening understanding of
animal culture brings novel perspectives to conservation. Proceedings of the Royal Society B 288: 20202718 Journal article
Feyrer, L.J., M. Stewart, J. Yeung,
C. Soulier and H. Whitehead. 2021. Origin and persistence of markings in a
long-term photo-identification dataset reveal the threat of entanglement for
endangered northern bottlenose whales (Hyperoodon ampullatus). Frontiers
in Marine Science 8: 620804. PDF
Hersh, T.A., S. Gero, L. Rendell and H.
Whitehead. 2021. Using identity calls to
detect structure in acoustic datasets. Methods in Ecology and Evolution 12: 1668-1678 Journal
article
Kowarski,K., S. Cerchio, H. Whitehead and H.
Moors-Murphy. 2021. Where, when, and why do western North Atlantic humpback
whales begin to sing? Bioacoustics 31:450-469. Journal article
Kowarski, K.A. and H. Moors-Murphy. 2021. A review of big data analysis methods
for baleen whale passive acoustic monitoring. Marine Mammal Science 37:652–673 PDF
McComb-Turbitt, S., J. Costa, H. Whitehead and M.
Auger-Méthé. 2021. Small-scale spatial distributions
of long-finned pilot whales change over time, but foraging hotspots are
consistent: significance for marine wildlife tourism management. Marine
Mammal Science 37:1196–1211 PDF
Whitehead, H., T.D. Smith and L. Rendell. 2021.
Adaptation of sperm whales to open-boat whalers: rapid social learning on a
large scale? Biology Letters 17: 20210030 Journal article
PDF
Eguiguren A., E. Pirotta, K. Boerder, M.
Cantor, G. Merlen and H. Whitehead. 2020 Historical and contemporary habitat
use of sperm whales around the Galápagos Archipelago: Implications for
conservation. Aquatic Conservation: Marine and Freshwater Ecosystems
2020: 1–16. PDF
Feyrer, L., S.T. Zhao, H. Whitehead and C.J.D.
Matthews 2020. Prolonged maternal investment in northern bottlenose whales
alters our understanding of beaked whale reproductive life history. PLoS ONE15: e0235114. PDF
Kobayashi, H., H. Whitehead and M. Amano. 2020.
Long-term associations among male sperm whales (Physeter macrocephalus).
PLoS ONE 15(12): e0244204 PDF
Kowarski, K.A., B.J. Gaudet, A.J. Cole, E.E. Maxner, S.P. Turner, S.B. Martin,
H.D. Johnson, and J.E. Moloney. 2020. Near real-time marine mammal monitoring
from gliders: Practical challenges, system development, and management
implications. Journal of the Acoustical Society of America 148(3):
1215-1230. PDF
Whitehead, H. 2020. Cultural specialization and
genetic diversity: killer whales and beyond. Journal of Theoretical Biology
490: 110164 PDF
Brakes, S.R.X. Dall, L.M. Aplin, S. Bearhop, E.L. Carroll, P. Ciucci, V. Fishlock, J.K.B. Ford,
E.C. Garland, S.A. Keith, P.K. McGregor, S.L. Mesnick, M.J. Noad, G. Notarbartolo di Sciara, M.M. Robbins, M.P. Simmonds, F.
Spina, A. Thornton, P.R. Wade, M.J. Whiting, J. Williams, L. Rendell, H.
Whitehead, A. Whiten and C. Rutz. 2019. Animal cultures matter for
conservation. Understanding the rich social lives of animals benefits international
conservation efforts. Science 363:
1032-1034. PDF
Cantor, M., S. Gero, H. Whitehead and L. Rendell. 2019. Sperm whale, the largest toothed
creature on Earth. In: Ethology and
behavioral ecology odontocetes. Edited by B. Würsig.
Springer: 261-280. Info
Clarke, E., L.J. Feyrer, H. Moors-Murphy and J.
Stanistreet. 2019. Click characteristics of northern bottlenose whales (Hyperoodon
ampullatus) and Sowerby's beaked whales (Mesoplodon bidens)
off eastern Canada. Journal of the Acoustical Society of America 146:
307-315. PDF
Eguiguren, A., E. Pirotta, M. Cantor, L. Rendell and H. Whitehead. 2019 Habitat
use of culturally distinct Galápagos sperm whale (Physeter macrocephalus) clans. Marine
Ecology Progress Series 609: 257-270. PDF
Einfeldt, A.L., D.N. Orbach and L.J. Feyrer, 2019. A
method for determining sex and chromosome copy number: sex-by-sequencing
reveals the first two species of marine mammals with XXY chromosome condition. Journal
of Mammalogy 5: 1671-1677. PDF
Feyrer, L.J., P. Bentzen, H. Whitehead, I.G. Paterson
and A. Einfeldt. 2019. Evolutionary impacts differ between two exploited
populations of northern bottlenose whale (Hyperoodon ampullatus). Ecology
and Evolution 9:
13567–13584. PDF
Hooker, S. K., N. Aguilar De Soto, R.W. Baird, E.L.
Carroll, D. Claridge, L. Feyrer, P.J.O. Miller, A. Onoufriou,
G. Schorr, E. Siegal and H. Whitehead. 2019. Future directions in research on
beaked whales. Frontiers in Marine
Science 5: 514. PDF
Konrad, C.M., T. Frasier, H. Whitehead and S. Gero.
2018. Kin selection and allocare in sperm whales. Behavioral Ecology 30: 194-201. PDF
Kowarski, K.A., H. Moors-Murphy, E.E. Maxner, and S. Cerchio. 2019. Western
North Atlantic humpback whale fall and spring acoustic repertoire: Insight into
onset and cessation of singing behavior. Journal of the Acoustical Society
of America 145: 2305-2316. PDF
Rendell, L., M. Cantor, S. Gero, H.
Whitehead and J. Mann. Causes and consequences of female centrality in cetacean
societies. 2019. Philosophical
Transactions of the Royal Society B 374: 20180066. PDF
Weiss, M.N., D.W. Franks, D.P. Croft and H. Whitehead. 2019. Measuring the complexity of social associations using mixture models. Behavioral Ecology and Sociobiology 73: 8. PDF
Whitehead, H., K.N. Laland, L.
Rendell, R. Thorogood and A. Whiten. 2019. The reach of gene-culture
coevolution in animals. Nature
Communications 10: 1038.
PDF
Beslin, W.A.M., H. Whitehead and S. Gero. Automatic
acoustic estimation of sperm whale body lengths achieved through machine
recognition of on-axis clicks. Journal of
the Acoustical Society of America 144: 3485-3495. PDF
Bøttcher, A., S. Gero, K. Beedholm, H. Whitehead and
P.T. Madsen. 2018. Variability of the
inter-pulse interval in sperm whale clicks with implications for size
estimation and individual identification. Journal
of the Acoustical Society of America 144: 365-374. PDF
Konrad, C.M., S. Gero, T. Frasier and H. Whitehead.
2018. Kinship influences sperm whale social organization within, but generally
not among, social units. Royal Society Open Science 5: 180914. PDF
Konrad, C.M., T. Frasier, L. Rendell, H. Whitehead and
S. Gero. 2018. Kinship and association do not explain vocal repertoire
variation among individual sperm whales or social units. Animal Behaviour 145: 131-140. PDF
Kowarski, K.A., C. Evers, H. Moors‐Murphy, B. Martin, and S.L. Denes. 2018. Singing through winter nights: Seasonal and diel occurrence of humpback whale (Megaptera novaeangliae) calls in and around the Gully MPA, offshore eastern Canada. Marine Mammal Science 34: 169-189. PDF
Vachon, F., H. Whitehead and T.R. Frasier. 2018. What factors shape genetic diversity in cetaceans? Ecology and Evolution 8: 1554-1572. PDF
Whitehead, H. 2018. Culture and social learning in
marine mammals. In Encyclopedia of Marine Mammals, 3rd ed. Edited by B Würsig, J.G.M. Thewissen and K.M.
Kovacs. Academic Press, San Diego: 232-234.
Whitehead, H. 2018. Sperm whale. In Encyclopedia of
Marine Mammals, 3rd ed. Edited by B Würsig,
J.G.M. Thewissen and K.M. Kovacs. Academic Press, San
Diego: 919-925.
Whitehead, H., and J.K.B. Ford. 2018. Consequences of
culturally-driven ecological specialization: killer whales and beyond. Journal of Theoretical Biology 456:
279-294. PDF
Wong, J.B. and M. Auger-Méthé.
2018. Using laser photogrammetry to measure long-finned pilot whales (Globicephala melas). Proceedings of the Nova Scotian Institute of
Science 49: 269-291. PDF
Würsig, B. and H. Whitehead. 2018. Aerial behavior. In Encyclopedia of
Marine Mammals, 3rd ed. Edited by B Würsig,
J.G.M. Thewissen and K.M. Kovacs. Academic Press, San
Diego: 6-10.
Augusto, J.F., T.R. Frasier and H. Whitehead. 2017. Social structure of long-finned pilot whales (Globicephala melas) off northern Cape Breton Island, Nova Scotia. Behaviour 154: 509-540. PDF
Cantor, M., A. Eguiguren,
G. Merlen and H. Whitehead. 2017. Galápagos sperm whales (Physeter macrocephalus): waxing and waning over three decades. Canadian
Journal of Zoology 95: 645–652. PDF
Cantor, M., M.M. Pires, F.M. Marquitti, R.L. Raimundo, E. Sebastián-González, P.P.
Coltri, S.I. Perez, D.R. Barneche, D.Y. Brandt, K.
Nunes and F.G. Daura-Jorge. 2017. Nestedness across
biological scales. PLOS ONE 12:
0171691. PDF
Konrad, C.M., A. Dupuis, S. Gero and T. Frasier. 2017
A sexing technique for highly degraded cetacean DNA. Aquatic Mammals 43: 655-660. PDF
Orbach, D.N., B. Hedrick, B. Würsig, S.L. Mesnick and P.L.R. Brennan. 2017. The
evolution of genital shape variation in female cetaceans. Evolution PDF
Orbach, D.N., D.A. Kelly, M. Solano
and P.L.R. Brennan PLR. 2017. Genital interactions during simulated copulation
among marine mammals.
Proceedings of the Royal Society B 284: 20171265. PDF
Whitehead, H. 2017. Gene-culture
coevolution in whales and dolphins. Proceedings
of the National Academy of Sciences (USA) 114: 7814-7821. PDF
Whitehead,
H. and L. Rendell. 2017. Deep
culture. In: Deep Thinkers. Edited by J.
Mann. Ivy Press, London: 122-143. Info
Whitehead, H., F. Vachon and T.R. Frasier. 2017. Cultural hitchhiking in the matrilineal whales, Behavior Genetics 47: 324-334. PDF
Zwamborn, E., and H. Whitehead. 2017. The baroque potheads: modification and
embellishment in repeated call sequences of long-finned pilot whales. Behaviour 154: 963-979. PDF
Atkins, S., M. Cantor, N. Pillay, G. Cliff, M. Keith and G.J. Parra. 2016. Net loss of endangered humpback dolphins: integrating residency, site fidelity, and bycatch in shark nets. Marine Ecology Progress Series 555: 249-260. PDF
Augusto, J.F., T.R. Frasier and H. Whitehead. 2016. Characterizing alloparental care in the pilot whale (Globicephala melas) population that summers off Cape Breton, Nova Scotia, Canada. Marine Mammal Science 33: 440-456. PDF
Beirão-Campos, L., M. Cantor, L. Flach and P.C. Simões-Lopes. 2016 Guiana dolphins form social modules in a large population with high ranging overlap and small demographic changes. Behavioral Ecology and Sociobiology 70: 1821-1830. PDF
Cantor, M., H. Whitehead, S. Gero and L. Rendell. 2016.
Cultural turnover among Galápagos sperm whales. Royal Society Open Science 3: 160615. PDF
Findlay, R., E. Gennari, M. Cantor and D.P. Tittensor.
2016. How solitary are white sharks: social interactions or just spatial
proximity? Behavioral Ecology and
Sociobiology 70: 1735-1744. PDF
Franks, D.W., S. Nattrass, L.J.N. Brent, H.
Whitehead, A.D. Foote, S. Mazzi, J.K.B. Ford, K.C. Balcomb, M.A. Cant and D. P.
Croft. 2016. The significance of post-reproductive lifespans in killer whales:
Reply to Robeck et al. Journal of Mammalogy 97: 906-909. PDF
Gero, S., A. Bøttcher, H. Whitehead and P. Madsen. 2016. Socially segregated, sympatric sperm whale clans in the Atlantic Ocean. Royal Society Open Science 3: 160061. PDF
Gero, S., and H. Whitehead. 2016. Critical decline of the
eastern Caribbean sperm whale population. PLOS
ONE 11:e0162019. PDF
Gero, S., H. Whitehead and L. Rendell. 2016. Individual, unit, and vocal clan level identity cues in sperm whale codas. Royal Society Open Science 3:150372. PDF
Rathinakumar, A., M. Cantor, K. Senthilkumar, P. Vimal, P. Kaliraj and G. Marimuthu. 2016. Social grooming among Indian short-nosed fruit bats. Behaviour 154: 37-63. PDF
Whitehead. H. 2016. Consensus movements by groups of sperm
whales. Marine Mammal Science 32:1402-1415. PDF
Yen, J.D.L., R.B. Cabral, M. Cantor, I. Hatton, S. Kortsch, J. Patrício and M. Yamamichi. 2016. Linking structure and function in food webs: maximization of different ecological functions generates distinct food web structures. Journal of Animal Ecology 85: 537–547. PDF
Zwamborn, E., and H. Whitehead.
2016. Repeated call sequences and behavioural context in long-finned pilot
whales off Cape Breton, Nova Scotia, Canada. Bioacoustics 26:
169-183. PDF
Au, W., J. Ford, L.M. Herman, P., Nachtigall, S.
Ridgway, J. Thomas, R. Wells, H. Whitehead, C.M. Johnson and D.L. Herzing.
2015. Visions of the future. In Dolphin
Communication & Cognition. Edited by C.M. Johnson and D.L.
Herzing. MIT Press, Cambridge, MA. Info
Cantor, M. and S. Gero. 2015. The missing metric: quantifying contributions
of reviewers. Royal Society Open Science
2: 140540. PDF
Cantor, M., L.G. Shoemaker, R.B. Cabral, C.O. Flores, M.Varga and H.
Whitehead. 2015. Multilevel animal societies can emerge from cultural
transmission. Nature Communications
6: 8091. PDF
Cantor, M. and H. Whitehead. 2015. How does social
behavior differ among sperm whale clans? Marine
Mammal Science 31: 1275-1290. PDF
Farine, D.R., and H. Whitehead. 2015. Constructing,
conducting and interpreting animal social network analysis. Journal of Animal Ecology 84: 1144–1163.
PDF
Gero, S., J. Gordon and H. Whitehead. 2015. Individualized social preferences and
long-term social fidelity between social units of sperm whales. Animal Behaviour
102: 15-23. PDF
Pires, M.M., M. Cantor, P.R. Guimarães, M. Aguiar,
S.F. Reis and P. Coltri. 2015. The network organization of protein interactions
in the spliceosome is reproduced by the simple rules of food-web models. Scientific Reports 5, 14865. PDF
Whitehead, H. 2015.
Life history evolution: What does a menopausal killer whale do? Current Biology 25: R225-226. PDF
Whitehead, H. and S. Gero. 2015. Conflicting rates of
increase in the sperm whale population of the eastern Caribbean: positive
observed rates do not reflect a healthy population. Endangered Species Research 27: 207-218. PDF
Whitehead, H. and R. James. 2015. Generalized
affiliation indices extract affiliations from social network data. Methods in Ecology and Evolution
6:836-844. PDF
Whitehead, H. and L. Rendell. 2015. The cultural lives of whales and dolphins.
University of Chicago Press. Info
Beirão, L., M. Cantor, L. Flach and C.A.B. Galdino. 2014. Performance of computer-assisted photographic matching of Guiana dolphins (Sotalia guianensis). Aquatic Mammals 40: 313-316. PDF
Cavechia, L., M. Cantor, A. Begossi and N. Peroni.
2014. Resource-use patterns in swidden farming communities: Implications for
resilience of cassava diversity. Human
Ecology 42: 605-616. PDF
Gero,
S., M. Milligan, C. Rinaldi, P. Francis, J. Gordon, C. Carlson, A. Steffen, P.
Tyack, P. Evans and H. Whitehead. 2014. Behavior and social structure of the
sperm whales of Dominica, West Indies. Marine
Mammal Science 30: 905-922. PDF
Kowarski, K.A., J.F. Augusto, T.R. Frasier and H. Whitehead. 2014. Effects of remote biopsy sampling on long-finned pilot whales (Globicephala melas) in Nova Scotia. Aquatic Mammals 40: 117-125. PDF
Lodi, L., M. Cantor, F.G. Daura-Jorge and C. Monteiro-Neto. 2014. A missing piece from a bigger puzzle: declining occurrence of a transient group of bottlenose dolphins off southeastern Brazil. Marine Ecology 35: 516-527. PDF
Moors-Murphy, H.B. 2014. Submarine canyons as important habitat for cetaceans, with special reference to the Gully: A review. Deep-Sea Research II 104: 6-19. PDF
Whitehead, H. and S. Gero. 2014. Using social structure to improve mortality estimates; an example with sperm whales. Methods in Ecology and Evolution 5: 27-36. PDF
Wong, S.N.P., and H. Whitehead. 2014. Seasonal occurrence of sperm whales (Physeter macrocephalus) around Kelvin Seamount in the Sargasso Sea in relation to oceanographic processes. Deep Sea Research I 91: 10-16. PDF
Augusto, J.F., T.R. Frasier and H. Whitehead. 2013.
Using photography to determine sex in pilot whales (Globicephala melas) is not possible: Males and females have similar
dorsal fins. Marine Mammal Science
29: 213-220. PDF
Cantor, M., and H. Whitehead. 2013. The interplay
between social networks and culture: theoretically and among whales and
dolphins. Philosophical Transactions of the Royal Society B 368. PDF
Gero, S., J. Gordon and H. Whitehead. 2013. Calves as
social hubs: dynamics of the social network within sperm whale units. Proceedings
of the Royal Society B 280: 20131113. PDF
Godde, S., L. Humbert, S.D. Côté, D. Réale, and H. Whitehead. 2013. Correcting for the impact of
gregariousness in social network analyses.
Animal Behaviour
85: 553-558. PDF
Gómez-Salazar, C., F. Trujillo and H. Whitehead. 2013.
Population size estimates of pink river dolphins (Inia geoffrensis) using mark-recapture
methods on photo-identification. Latin
American Journal of Aquatic Mammals 9: 40-46. PDF
O’Brien, K. and H. Whitehead. 2013. Population analysis
of Endangered northern bottlenose whales on the Scotian Shelf seven years after
the establishment of a Marine Protected Area.
Endangered Species Research
21: 273–284. PDF
Whitehead, H. and I.D. Jonsen. 2013. Inferring animal
densities from tracking data using Markov chains. PLoS ONE 8(4): e60901. PDF
Whitehead, H. 2013. Trends in cetacean abundance in
the Gully submarine canyon, 1988–2011, highlight a 21% per year increase in
Sowerby’s beaked whales (Mesoplodon bidens). Canadian
Journal of Zoology 91: 141-148. PDF
Publications
2012
Gomez-Salazar, C., M. Coll, and H. Whitehead. 2012.
River dolphins as indicators of ecosystem degradation in large tropical rivers.
Ecological Indicators 23: 19-26. PDF
Gomez-Salazar, C., F. Trujillo, M. Portocarrero-Aya
and H. Whitehead. 2012. Population, density estimates and conservation of river
dolphins (Inia and Sotalia) in the
Amazon and Orinoco river basins. Marine
Mammal Science 28:124-153. PDF
Gomez-Salazar, C., F. Trujillo, and H. Whitehead.
2012. Ecological factors influencing group sizes of river dolphins (Inia geoffrensis
and Sotalia fluviatilis). Marine Mammal Science 33: 1142-1164. PDF
Rendell, L., S.L. Mesnick, M.L. Dalebout, J.
Burtenshaw and H. Whitehead. 2012. Can
genetic differences explain vocal dialect variation in sperm whales, Physeter macrocephalus? Behavior Genetics 42: 332-343. PDF
Senigaglia, V. and H. Whitehead. 2012. Synchronous breathing by pilot whales. Marine Mammal Science 28: 213-219. PDF
Whitehead, H., R. Antunes, S. Gero, S.N.P. Wong, D.
Engelhaupt and L. Rendell. 2012. Multilevel societies of female sperm whales (Physeter macrocephalus) in the Atlantic
and Pacific: why are they so different? International
Journal of Primatology 33: 1142-1164. PDF
Whitehead, H. and S.K. Hooker. 2012. Uncertain status
of the northern bottlenose whale (Hyperoodon
ampullatus): population fragmentation, legacy of whaling, and current
threats. Endangered Species Research
19: 47-61. PDF
Whitehead, H., and D. Lusseau. 2012. Animal social
networks as substrate for cultural behavioural
diversity. Journal of Theoretical Biology
294: 19-28. PDF
Antunes, R., T. Schulz, S. Gero, H. Whitehead, J.
Gordon and L. Rendell. 2011. Individually distinctive acoustic features in
sperm whale codas. Animal Behaviour 81: 723-730. PDF
Auger-Méthé, M., M. Marcoux,
H. Whitehead. 2011. Computer-assisted photo-identification of narwhals. Arctic 64: 342-352. PDF
Caro, T., K. Beeman, T. Stankowich
and H. Whitehead. 2011. The functional significance of colouration
in cetaceans. Evolutionary Ecology
25: 1231-1245. PDF
Gomez-Salazar, C., F. Trujillo, and H. Whitehead.
2011. Photo-identification: a promising and noninvasive tool to study pink
river dolphins (Inia geoffrensis).
Aquatic Mammals 37: 473-485. PDF
Horrocks, J., D.C. Hamilton and H. Whitehead. 2011. A
likelihood approach to estimating animal density from binary acoustic
transects. Biometrics 67: 681-690. PDF
Schulz, T.M., H. Whitehead, S. Gero and L. Rendell.
2011. Individual vocal production in a sperm whale (Physeter macrocephalus) social unit. Marine Mammal Science 27: 149-166. PDF
Whitehead, H.
2011. The cultures of whales and
dolphins. In: Whales and dolphins:
cognition, culture, conservation and human perceptions. Edited by P. Brakes and M. P. Simmonds. Earthscan, London, U.K. Info
Auger‑Méthé, M., Marcoux, M. and H. Whitehead. 2010. Nicks and notches of the dorsal ridge: Promising mark types for the photo-identification of narwhals. Marine Mammal Science 26: 663-678. PDF
Wade, L., H. Whitehead and L. Weilgart. 2010.
Conflict of interest in research on anthropogenic noise and marine
mammals: does funding bias conclusions? Marine
Policy 34: 320-327. PDF
Whitehead, H.
2010. Conserving and managing
animals that learn socially and share cultures. Learning and Behavior 38: 329-336. PDF
Whitehead,
H., O’Brien, K. and B. Worm. 2010.
Diversity of deep-water cetaceans and primary productivity. Marine
Ecology Progress Series 408: 1-5. PDF
Whitehead, H. and S. Van Parijs. 2010. Studying
marine mammal social systems. In: Marine mammal ecology and conservation: a
handbook of techniques. Edited by I.
Boyd, D. Bowen and S. Iverson. Oxford
University Press, Oxford, U.K. PDF
Info
Bejder, L., A. Samuels, H. Whitehead, H. Finn and S.
Allen. 2009. Use and misuse of habituation, sensitisation
and tolerance to describe wildlife responses to anthropogenic stimuli in impact
assessment research. Marine Ecology
Progress Series 395: 177-185. PDF
Engelhaupt,D., A.R. Hoelzel, C. Nicholson, A. Frantzis, S. Mesnick, S. Gero, H.
Whitehead, L. Rendell, P. Miller, R. De Stefanis, A. Cañadas, S. Airoldi and
A.A. Mignucci-Giannoni. 2009. Female philopatry in coastal basins and male
dispersion across the North Atlantic in a highly mobile marine species, the
sperm whale (Physeter macrocephalus).
Molecular Ecology 18: 4193–4205. PDF
Gero,
S., D. Engelhaupt, L. Rendell and H. Whitehead.
2009. Who Cares? Between-group
variation in alloparental care-giving in sperm whales. Behavioral
Ecology 20: 838-843. PDF
Nemiroff, L., and H. Whitehead. 2009. Structural characteristics of pulsed calls of long-finned pilot whales Globicephala melas. Bioacoustics 19: 67-92. PDF
Schulz,
T., H. Whitehead, and L. Rendell. 2009. Off-axis effects on the multi-pulse
structure of sperm whale coda clicks. Journal of the Acoustical Society of America 125: 1768-1773. PDF
Whitehead, H., and P.J. Richerson. 2009. The evolution of conformist social learning can cause population collapse in realistically variable environments. Evolution and Human Behavior 30: 261-273. PDF
Whitehead, H. 2009. Estimating abundance from one‑dimensional passive acoustic surveys. Journal of Wildlife Management 73: 1000-1009. PDF
Whitehead, H. 2009. SOCPROG programs: analyzing animal
social structures. Behavioral Ecology and
Sociobiology 63:765-778. PDF
Whitehead, H. 2009. How might we study culture? A
perspective from the ocean. In: The
question of animal culture. Edited
by R. N. Laland, and B. Galef. Harvard University Press, Cambridge, MA.:
125-152.
Gero, S., D. Engelhaupt, and H. Whitehead. 2008.
Heterogeneous social associations within a sperm whale, Physeter macrocephalus, unit reflect pairwise relatedness. Behavioral Ecology and Sociobiology 63:
143-151. PDF
Higham, J.E.S.
and D. Lusseau. 2008. Slaughtering the goose that lays the golden egg. Are
whaling and whalewatching mutually exclusive? Current
Issues in Tourism 11: 63-74.
Hooker, S. K., T. L. Metcalfe, C. D. Metcalfe, C. M.
Angell, J. Y. Wilson, M. J. Moore and H. Whitehead. 2008.
Changes in persistent contaminant concentration and CYP1A1 protein
expression in biopsy samples from northern bottlenose whales, Hyperoodon ampullatus, following the
onset of nearby oil and gas development.
Environmental Pollution 152:
205-216. PDF
Lusseau, D., H. Whitehead and S. Gero. 2008. Incorporating
uncertainty into the study of animal social networks. Animal Behaviour 75: 1809-1815. PDF
Marino, L., C. Butti, R. C. Connor, R. E. Fordyce, L.
M. Herman, P. R. Hof, L. Lefebvre, D. Lusseau, B. McCowan, E. A. Nimchinsky, A. A. Pack, J. S. Reidenberg,
D. Reiss, L. Rendell, M. D. Uhen, E. Van der Gucht
and H. Whitehead. 2008. A claim in search of evidence: Reply to
Manger's thermogenesis hypothesis of cetacean brain structure. Biological
Reviews 83: 417-440. PDF
Schulz, T., H. Whitehead, S. Gero, and L. Rendell.
2008. Overlapping and matching of codas in vocal interactions between sperm
whales: insights into communication function. Animal Behaviour 76:1977-1988. PDF
Smith, T.D., R.R. Reeves,
E.A. Josephson, J.N. Lund and H. Whitehead.
2008. Sperm whale catches and
encounter rates during the 19th and 20th centuries: an apparent paradox. In: Oceans
past: management insights from the history of marine animal populations. Edited by D.J. Starkey, P. Holm, and M.
Barnard. James & James / Earthscan,
London, U.K.: 149-173. PDF
Whitehead, H.
2008. Precision and power in the
analysis of social structure using associations. Animal Behaviour 75: 1093-1099. PDF
Whitehead, H.
2008. Analyzing animal societies: quantitative methods for vertebrate social
analysis. University of Chicago Press, Chicago. Info
Whitehead, H. 2008.
Social and cultural evolution in the ocean. Convergences and contrasts
with terrestrial systems. In: The Deep Structure of Biology. Edited by S. Conway-Morris. Templeton Foundation Press, West
Conshohocken, Pennsylvania: 143-160.
Whitehead, H., B. McGill and B. Worm. 2008.
Diversity of deep-water cetaceans in relation to temperature:
implications for ocean warming. Ecology Letters 11: 1198-1207. PDF
Whitehead, H., A. Coakes, N.
Jaquet and S. Lusseau. 2008. Movements of sperm whales in the tropical Pacific.
Marine Ecology Progress Series 361:
291-300. PDF
Publications 2007
Auger-Méthé, M. and H. Whitehead. 2007. The use of natural
markings in studies of long-finned pilot whale (Globicephala melas). Marine
Mammal Science 23: 77-93. PDF
Gero, S. and H.
Whitehead. 2007. Suckling behavior in sperm whales calves: observations and
hypotheses. Marine Mammal Science 23: 398-413. PDF
Gero, S., J. Gordon, C. Carlson, P. Evans and H.
Whitehead. 2007. Population estimate and
inter-island movement of sperm whales, Physeter
macrocephalus, in the Eastern Caribbean sea. Journal of
Cetacean Research and Management 9: 143-150. PDF
Higham, J.E.S.
and Lusseau D. 2007. Urgent need for empirical research into whaling and whalewatching. Conservation Biology 21: 554-558. PDF
Higham, J.E.S.
and Lusseau D. 2007. Defining critical habitats: the spatio-ecological
approach to managing tourist-wildlife interactions. In: Critical Issues in
Ecotourism: Understanding a complex tourism phenomenon. Edited by J.E.S. Higham.
Elsevier Butterworth-Heinemann Publications, Oxford, UK: 256-269.
Higham, J.E.S.
and D. Lusseau. 2007. The spatio-ecological approach
to managing wildlife-tourist interactions. In: Marine Wildlife and Tourism
Management Edited by J.E.S. Higham, and M. Lück. CAB International
Publishing, Oxford, UK.
Lusseau, D.
2007. Understanding the impacts of noise on marine mammals. In: Marine
Wildlife and Tourism Management Edited by J.E.S. Higham, and M. Lück. CAB
International Publishing, Oxford, UK.
Lusseau, D.
2007. Why are male social relationships complex in the Doubtful Sound
bottlenose dolphin population? PLoS ONE
2(4): e348. doi10.1371/journal.pone.0000348 PDF
Lusseau, D.
2007. Evidence for social role in a dolphin social network. Evolutionary
Ecology 21(3). PDF
Marcoux, M., L. Rendell, and H. Whitehead. 2007.
Indications of fitness differences among vocal clans of sperm
whales. Behavioural Ecology and Sociobiology 61: 1093-1098. PDF
Marcoux, M., H.
Whitehead, and L. Rendell. 2007. Sperm whale feeding variation by location,
year, social group and clan: Evidence from stable isotopes. Marine Ecology
Progress Series 333:309-314. PDF
Marino, L., R.
C. Connor, R. E. Fordyce, L. M. Herman, P. R. Hof, L. Lefebvre, D. Lusseau, B.
McCowan, E. A. Nimchinsky, A. A. Pack, L. Rendell, J.
S. Reidenberg, D. Reiss, M. D. Uhen,
E. Van der Gucht and H. Whitehead.
2007. Cetaceans have complex
brains for complex cognition. PDF
Weilgart, L. 2007. The impacts of
anthropogenic ocean noise on cetaceans and implications for management. Canadian
Journal of Zoology 85(11):1091–1116. PDF
Whitehead, H.
2007. Learning, climate and the evolution of cultural capacity. Journal of
Theoretical Biology 245: 341-350. PDF
Whitehead, H. 2007.
Selection of models of lagged identification rates and lagged association rates
using AIC and QAIC. Communications in
Statistics–Simulation and Computation 36:1233-1246.
PDF
Publications 2006
Bejder, L., A.
Samuels, H. Whitehead, and N. Gales. 2006. Interpreting short-term behavioural responses to disturbance within a longitudinal
perspective. Animal Behaviour 72:1149-1158. PDF
Bejder, L., A.
Samuels, H. Whitehead, N. Gales, J. Mann, R. Connor, M. Heithaus, J.
Watson-Capps, and C. Flaherty. 2006. Decline in relative abundance of
bottlenose dolphins exposed to long-term disturbance. Conservation Biology 20:
1791-1798. PDF
Dalebout, M.
L., D. E. Ruzzante, H. Whitehead, and N. I. Øien. 2006. Nuclear and
mitochondrial markers reveal distinctiveness of a small population of
bottlenose whales (Hyperoodon ampullatus) in the western North Atlantic.
Molecular Ecology 15:3115-3129. PDF
Ferrer i Cancho, R., and D. Lusseau.
2006. Long-range correlation in the surface behaviour
of dolphins. Europhysics Letters 74(6):
1095-1101. PDF
Lusseau, D.
2006. The short-term behavioral reactions of bottlenose dolphins to
interactions with boats in doubtful sound, New Zealand. Marine Mammal
Science 22:802-818. PDF
Lusseau, D., E.
Slooten, and R.J. Currey. 2006. Unsustainable dolphin watching activities in
Fiordland, New Zealand. Tourism in Marine Environments 3(2) PDF
Lusseau, D.
2006. Why do dolphins jump? Interpreting the behavioural
repertoire of bottlenose dolphins in Doubtful Sound, New Zealand. Behavioural Processes PDF
Marcoux, M., H.
Whitehead, and L. Rendell. 2006. Coda vocalizations recorded in breeding areas
are almost entirely produced by mature female sperm whales (Physeter
macrocephalus). Canadian Journal of Zoology 84: 609-614. PDF
Schulz, T., L. Rendell, H.
Whitehead. 2006. A remotely-piloted acoustic array for studying sperm whale
vocal behaviour. Canadian
Acoustics 34: 54-55.
Weinrich, M.
T., H. Rosenbaum, C. S. Baker, A. L. Blackmer, and H. Whitehead. 2006. The
influence of maternal lineages on social affiliations among humpback whales (Megaptera
novaeangliae) on their feeding grounds in the Southern Gulf of Maine. Journal
of Heredity 97: 226-234. PDF
Whitehead, H.
2006. Sperm whales in ocean ecosystems. In: Whales, whaling and ocean
ecosystems. Edited by J. A. Estes, D. P. DeMaster, D. F. Doak, T. M.
Williams, and R. L. Brownell. University of California Press, Berkeley, CA:
324-334.
Williams, R.,
and D. Lusseau. 2006. A killer whale social network is vulnerable to targeted
removals. Biology Letters PDF
Williams, R.,
D. Lusseau and P.S. Hammond P.S. 2006. Potential energetic cost to killer
whales of disturbance by vessels and the role of a marine protected area. Biological
Conservation PDF
Publications 2005
Beekmans, B. W. P. M., H. Whitehead, R. Huele, L.
Steiner, and A. G. Steenbeek. 2005. Comparison of two
computer-assisted photo-identification methods applied to sperm whales (Physeter
macrocephalus). Aquatic Mammals 31: 243-247. PDF
Connor, R., and
H. Whitehead. 2005. Alliances II. rates of encounter during resource
utilization: a general model of intrasexual alliance formation in
fission-fusion societies. Animal Behaviour 69:
127-132. PDF
Dalebout, M.
L., K. M. Robertson, A. Frantzis, D. Engelhaupt, A. A. Mignucci-Giannoni, R.J.
Rosario-Delestre, and C. S. Baker. 2005. Worldwide
structure of mtDNA diversity among Cuvier's beaked
whales (Ziphius cavirostris); implications for
threatened populations. Molecular Ecology PDF
Endo, T., K.
Haraguchi, Y. Hotta, Y. Hisamichi, S. Lavery, M. L.
Dalebout, and C. S. Baker. 2005. Total mercury, methyl mercury, and selenium
levels in the red meat of small cetaceans sold for human consumption in Japan. Environmental
Science and Technology 39: 5703-5708. PDF
Gero, S., L. Bejder,
H. Whitehead, J. Mann and R. C. Connor. 2005. Behaviourally
specific preferred associations in bottlenose dolphins, Tursiops sp.
Canadian Journal of Zoology 83:1566-1573 PDF
McAuliffe, K.
and H. Whitehead. 2005. Eusociality, menopause and information in matrilineal
whales. Trends in Ecology and Evolution 20: 650. PDF
Rendell, L. and
H. Whitehead. 2005. Coda playbacks to sperm whales in Chilean waters. Marine
Mammal Science 21: 106-115.
Rendell, L. E.,
and H. Whitehead. 2005. Spatial and temporal variation in sperm whale coda vocalisations: Stable usage and local dialects. Animal Behaviour 70: 191-198. PDF
Rendell, L., H.
Whitehead and A. Coakes. 2005. Do breeding male sperm
whales show preferences among vocal clans of females? Marine Mammal Science
21: 116-120.
Weilgart, L, H. Whitehead, L. Rendell and J. Calambokidis.
2005. Signal-to-noise: funding structure versus ethics as a solution to
conflict-of-interest. Marine Mammal Science 20: 898-899.PDF
Email exchange between
representative of operational US Navy and official of Office of Naval Research (Notarized
authentication of email exchange)
Whitehead, H.
2005. Genetic diversity in the matrilineal whales: models of cultural
hitchhiking and group-specific non-heritable demographic variation. Marine
Mammal Science 21: 58-79.PDF
Whitehead, H.,
L. Bejder, and A. C. Ottensmeyer. 2005. Testing
association patterns: issues arising and extensions. Animal Behaviour 69: e1-e6. PDF
Whitehead, H.,
and R. Connor. 2005. Alliances I: how large should alliances be? Animal Behaviour 69: 117-126. PDF
Whitehead, H., and
R. Reeves. 2005. Killer whales and whaling: the scavenging hypothesis.
Biology Letters PDF Appendix
Whitehead, H.,
and T. Wimmer. 2005. Heterogeneity and the mark-recapture assessment of the
Scotian Shelf population of northern bottlenose whales (Hyperoodon
ampullatus). Canadian Journal of Fisheries and Aquatic Sciences
63:2573-2585. PDF
Wimmer, T., and
H. Whitehead. 2005. Movements and distribution of northern bottlenose whales,Hyperoodon ampullatus, on the Scotian
Slope and in adjacent waters. Canadian Journal of Zoology 82: 1782-1794.
PDF
Publications 2004
Coakes, A.K, and H. Whitehead. 2004. Social structure and mating system of
sperm whales off northern Chile. Canadian Journal of Zoology 82:
1360-1369. PDF
Dalebout, M.L.,
C.S. Baker, J. G. Mead, V. G. Cockcroft and T. K. Yamada. 2004. A comprehensive
and validated molecular taxonomy of beaked whales, family Ziphiidae. Journal
of Heredity 95: 459-473. PDF
Dalebout, M.L.,
K. G. Russell, M. J. Little and P. Ensor. 2004. Observations of live Gray’s
beaked whales (Mesoplodon grayi) in Mahurangi Harbour, North Island,
New Zealand, with a summary of at-sea sightings. Journal of the Royal
Society of New Zealand 34: 347-356. PDF
Durrell, J. L.,
I. A. Sneddon, N. E. O’Connell, and H. Whitehead. 2004. Do pigs form
preferential associations? Applied Animal Behaviour
Science 89: 41-52. PDF
Rendell, L. E.,
and H. Whitehead. 2004. Do sperm whales share coda vocalizations? Insights into
coda usage from acoustic size measurement. Animal Behaviour
67: 865-874. PDF
Rendell, L., H.
Whitehead, and R. Escribano. 2004. Sperm whale habitat use and foraging success
off northern Chile: evidence of ecological links between coastal and pelagic
systems. Marine Ecology Progress Series 275: 289-295. PDF
Samuels, A. and
L. Bejder. 2004. Chronic interactions between humans and wild bottlenose
dolphins (Tursiops truncatus) near Panama City Beach, Florida. Journal
of Cetacean Research and Management 6: 69-77.
Vonhof, M. J.,
H. Whitehead, and M. B. Fenton. 2004. Analysis of Spix's disk-winged bat
association patterns and roosting home ranges reveal a novel social structure
among bats. Animal Behaviour 68: 507-521. PDF
Whitehead, H. 2004.
The group strikes back: follow protocols for behavioral research on cetaceans. Marine
Mammal Science 20: 304-310. PDF
Whitehead, H.
2004. Whale culture and conservation. In: Encyclopedia of animal behavior, Vol.
1. Edited by M. Bekoff. Greenwood Press, Wesport, Connecticut: 466-468.
Whitehead, H., and
L. Rendell. 2004. Movements, habitat use and feeding success of cultural clans
of South Pacific sperm whales. Journal of Animal Ecology 73:190-196. PDF
Whitehead, H.,
L. Rendell, R. W. Osborne, and B. Würsig. 2004.
Culture and conservation of non-humans with reference to whales and dolphins:
review and new directions. Biological Conservation 120: 431–441. PDF
Publications 2003
Allen, S. and
L. Bejder. 2003. Southern Right Whale (Eubalaena
australis) sightings on the Australian coast and the increasing potential
for entanglement. Pacific Conservation Biology 9: 228-233.
Bejder, L. and
A. Samuels. 2003. Evaluating impacts of nature-based tourism on cetaceans. In:
N. Gales, M. Hindell, R. Kirkwood (eds.) Marine Mammals: Fisheries, Tourism and
Management Issues. CSIRO Publishing. 480 pp.
Ottensmeyer, C. A., and H. Whitehead. 2003. Behavioural
evidence for social units in long-finned pilot whales. Canadian Journal of
Zoology 81:1327-1338. PDF
Rendell, L. E.,
and H. Whitehead. 2003. Comparing repertoires of sperm whale codas: a multiple
methods approach. Bioacoustics 14:59-79. PDF
Rendell, L.
& Whitehead, H. 2003. Vocal clans in sperm whales (Physeter
macrocephalus).Proceedings of the Royal Society: Biological Sciences.
270:225-231. PDF
Samuels A.,
Bejder, L., Constantine R., and Heinrich, S. 2003. A review of swimming with
wild cetaceans with a specific focus on the Southern Hemisphere. In: N. Gales,
M. Hindell, R. Kirkwood (eds.) Marine Mammals: Fisheries, Tourism and
Management Issues. CSIRO Publishing. 480 pp.
Weilgart, L. 2003. Acoustic smog. In: Between species; celebrating the
dolphin-human bond. Edited by T. Frohoff and B.
Peterson. Sierra Club Books, San Francisco:230-238.
Whitehead, H.,
C. D. Macleod, and P. Rodhouse. 2003. Differences in niche breadth among some teuthivorous mesopelagic marine mammals. Marine Mammal
Science 19: 400-406. PDF
Whitehead, H.
2003. Sperm whale societies; social evolution in the ocean. Chicago:
University of Chicago Press. 431pp .Info
Whitehead, H.
2003. Society and culture in the deep and open ocean: the sperm whale. In:Animal social complexity. Intelligence,
culture and individualized societies, ed. F. B. M. de Waal, and P. L.
Tyack. Cambridge, Massachusetts: Harvard University Press. pp. 444-464.
Publications 2002
Baird, R. W.
2002. Killer whales of the world: natural history and conservation.
Voyageur Press, Stillwater, MN
Baird, R. W.
2002. False killer whale Pseudorca crassidens. In Encyclopedia Of Marine
Mammals. Edited by W.F. Perrin, B. Wursig and J.G.M. Thewissen. Academic Press, San Diego
Baird, R. W.
2002. Risso's dolphin Grampus griseus In Encyclopedia of Marine Mammals.
Edited by W.F. Perrin, B. Wursig and J.G.M. Thewissen. Academic Press, San Diego
Baird, R. W.,
P. J. Stacey, D. A. Duffus and K. M. Langelier. 2002. An evaluation of gray
whale (Eschrichtius robustus) mortality
incidental to fishing operations in British Columbia, Canada. Journal of
Cetacean Research and Management 4: 289-296. PDF
Baird, R. W.,
J. F. Borsani, M. B. Hanson, and P. L. Tyack. 2002.
Diving and night-time behaviour of long-finned pilot
whales in the Ligurian Sea. Marine Ecology Progress Series 237: 301-305.
PDF
Bejder, L. & B.
K. Hall. 2002. Limbs in whales and limblessness in other
vertebrates: mechanisms of evolutionary and developmental transformation and
loss. Evolution and Development 4(6) : 445-458.
Berrow, S.D.,
B. McHugh, D. Glynn, E. McGovern, K.M. Parsons, R.W. Baird and S.K. Hooker.
2002. Organochlorine concentrations in resident bottlenose dolphins (Tursiops
truncatus) in the Shannon estuary, Ireland. Marine Pollution Bulletin 44:
1296-1313. PDF
Dillon, M.C.,
K.R. Richard, H. Whitehead and J.M. Wright. 2002. Social and population genetic
structure of sperm whales assessed by molecular genetic analyses. In: Molecular
and cell biology of marine mammals Edited by C. Pfeiffer. Krieger, Malabar,
Florida: 43-52.
Goold, J., H.
Whitehead, and J. Reid. 2002. North Atlantic sperm whale strandings on the
coastlines of the British Isles and eastern Canada. Canadian Field
Naturalist. 116:371-388.
Hooker, S.K.,
H. Whitehead, and S. Gowans. 2002. Ecosystem consideration in conservation
planning: energy demand of foraging bottlenose whales (Hyperoodon ampullatus)
in a marine protected area. Biological Conservation 104: 51-58. Abstract
Hooker, S.K.,
and H. Whitehead. 2002. Click characteristics of northern bottlenose whales (Hyperoodon
ampullatus). Marine Mammal Science 18: 69-80. Abstract
Hooker, S.K.,
H. Whitehead, S. Gowans, and R.W. Baird. 2002. Fluctuations in distribution and
patterns of individual range use of northern bottlenose whales. Marine
Ecology Progress Series225: 287-297. Abstract
Letteval, E., C. Richter, N. Jaquet, E. Slooten, S. Dawson, H. Whitehead, J.
Christal, and P. McCall Howard. 2002. Social structure and residency in
aggregations of male sperm whales.Canadian
Journal of Zoology 80: 1189-1196. PDF
Whitehead, H.
2002. Culture in whales and dolphins. In: Encyclopedia of marine mammals Edited
by W.F. Perrin, B. Würsig and H.G.M. Thewissen. Academic Press, San Diego
Whitehead, H. 2002.
Breaching. In: Encyclopedia of marine mammals Edited by W.F. Perrin, B. Würsig and H.G.M. Thewissen.
Academic Press, San Diego
Whitehead, H.
2002. The sperm whale. In: Encyclopedia of marine mammals Edited by W.F.
Perrin, B. Würsig and H.G.M. Thewissen.
Academic Press, San Diego
Whitehead, H.
2002. Culture in whales and dolphins. In: Encyclopedia of marine mammals Edited
by W.F. Perrin, B. Würsig and H.G.M. Thewissen. Academic Press, San Diego
Whitehead, H.
2002. Estimates of the current global population size and historical trajectory
for sperm whales. Marine Ecology Progress Series 242: 295-304. PDF
Whitehead, H.,
P. J. Richerson, and R. Boyd. 2002. Cultural selection and genetic diversity in
humans. Selection 3: 115-125. PDF
Publications 2001
Baird, R.W.
2001. Status of killer whales, Orcinus orca, in Canada. Canadian
Field-Naturalist115:676-701.
Baird, R.W.
2001. Status of harbour seals, Phoca
vitulina, in Canada. Canadian Field-Naturalist115:
663-675.
Baird, R.W.,
A.D. Ligon, S.K. Hooker, and A.M. Gorgone. 2001. Sub-surface and night-time behaviour of pantropical spotted dolphins in Hawaii. Canadian
Journal of Zoology 79: 988-996.Abstract
Bejder, L. and
Dawson, S. 2001 Abundance, residency, and habitat utilisation
of Hector's dolphins (Cephalorhynchus hectori) in Porpoise Bay, New Zealand. New Zealand
Journal of Marine and Freshwater Research: 35: 277-287. Abstract
Christal, J.,
and H. Whitehead. 2001. Social affiliations within sperm whale (Physeter
macrocephalus) groups. Ethology 107: 323-340.
PDF
Dalebout, M.L.,
S.K. Hooker, and I. Christensen. 2001. Genetic diversity and population
structure among northern bottlenose whales, Hyperoodon ampullatus, in
the western North Atlantic Ocean. Canadian Journal of Zoology 79:
478-484. Abstract
Gowans, S., H.
Whitehead, and S.K. Hooker. 2001. Social organization in northern bottlenose
whales (Hyperoodon ampullatus): not driven by deep water foraging? Animal
Behaviour 62: 369-377. PDF
Gowans, S. and
H. Whitehead. 2001. Photographic identification of northern bottlenose whales (Hyperoodon
ampullatus): sources of heterogeneity from natural marks. Marine Mammal
Science 17: 76-93. PDF
Hooker, S.K.,
and R.W. Baird. 2001. Diving and ranging behaviour of
odontocetes: a methodological review and critique. Mammal Review 31:
81-105. Abstract
Hooker, S.K.,
R.W. Baird, S. Al-Omari, S. Gowans and H. Whitehead. 2001. Behavioral reactions
of northern bottlenose whales to biopsy darting and tag attachment procedures. Fishery
Bulletin, U.S. 99: 303-308. Abstract
Hooker, S.K.,
S.J. Iverson, P. Ostrom, and S.C. Smith. 2001. Diet of northern bottlenose
whales inferred from fatty-acid and stable-isotope analyses of biopsy samples.Canadian Journal of Zoology
79:1442-1454. Abstract
Rendell, L. and
Whitehead, H. 2001. Culture in whales and dolphins. Behavioral and Brain
Sciences, 24(2): 309-382 PDF
Smith, S.C. and
H. Whitehead. 2001. Reply to R. Clarke and Paliza's
comment: "The food of sperm whales in the Southeast Pacific". Marine
Mammal Science 17: 430-431.
Whitehead, H.
2001. Analysis of animal movement using opportunistic individual
identifications: application to sperm whales. Ecology 82: 1417-1432. PDF
Whitehead, H.
2001. Direct estimation of within-group heterogeneity in photoidentification of
sperm whales. Marine Mammal Science 17:718-728. PDF
Publications 2000
Baird, R.W.,
and H. Whitehead. 2000. Social organization of mammal-eating killer whales:
group stability and dispersal patterns. Canadian Journal of Zoology
78:2096-2105. PDF
Baird, R.W.,
and S.K. Hooker. 2000. Ingestion of plastic and unusual prey by a juvenile harbour porpoise. Marine Pollution Bulletin 40:719-720.
PDF
Gowans, S., H.
Whitehead, J. Arch, and S.K. Hooker. 2000. Population size and residency
patterns of northern bottlenose whales (Hyperoodon ampullatus) using the
Gully. Journal of Cetacean Research and Management 2: 201-210. Abstract
Gowans, S.,
M.L. Dalebout, S.K. Hooker, and H. Whitehead. 2000. Reliability of photographic
and molecular techniques for sexing northern bottlenose whales (Hyperoodon
ampullatus). Canadian Journal of Zoology 78: 1224-1229. Abstract
Lucas, Z.N.,
and S.K. Hooker. 2000. Cetacean strandings on Sable Island, Nova Scotia,
1970-1998. Canadian Field-Naturalist 114: 45-61 Abstract
Mann, J., R.C.
Connor, P.L. Tyack, and H. Whitehead (Editors). 2000. Cetacean Societies.
Field studies of dolphins and whales. Chicago University Press. 432 pp More
information
∙ Baird, R.W.
The killer whale - foraging specializations and group hunting. Introduction
∙ Connor, R.C.,
J. Mann, P. Tyack and H. Whitehead. Introduction: the social lives of whales
and dolphins.
∙ Tyack, P., Connor,
R.C., J. Mann, and H. Whitehead. The future of behavioral research on wild
cetaceans.
∙ Whitehead, H.
and L. Weilgart. The sperm whale: social females and
roving males.
∙ Whitehead, H.
and J. Mann. Female reproductive strategies of cetaceans.
∙ Whitehead,
H., J. Christal and P. Tyack. Studying cetacean social structure in space and
time: innovative techniques.
∙ Whitehead,
H., R.R. Reeves and P. Tyack. Science and the conservation, protection and
management of wild cetaceans.
Smith, S.C.,
and H. Whitehead. 2000. The diet of Galapagos sperm whales Physeter
macrocephalus as indicated by fecal sample analysis. Marine Mammal
Science 16: 315-325. PDF
Whitehead, H.
2000. Density-dependent habitat selection and the modeling of sperm whale (Physeter
macrocephalus) exploitation. Canadian Journal of Fisheries and Aquatic
Sciences 57: 223-230. PDF
Publications 1999
Bejder, L.,
Dawson, S. and Harraway, J. 1999. Responses by Hector's dolphins to boats and
swimmers in Porpoise Bay, New Zealand. Marine Mammal Science:15(3):
738-750. Abstract
Dufault S, H.
Whitehead, and M. Dillon. 1999. An examination of the current knowledge on the
stock structure of sperm whales (Physeter macrocephalus) worldwide. Journal
of Cetacean Research and Management 1: 1-10. PDF
Gowans, S., and
L. Rendell. 1999. Head-butting in northern bottlenose whales (Hyperoodon
ampullatus): a possible function for big heads? Marine Mammal Science 15:
1342-1350. Abstract
Hooker, S.K.,
H. Whitehead and S. Gowans. 1999. Marine protected area design and the spatial
and temporal distribution of cetaceans in a submarine canyon. Conservation
Biology 13: 592-602.Abstract
Hooker, S.K.,
and R.W. Baird. 1999. Deep-diving behaviour of the
northern bottlenose whale, Hyperoodon ampullatus (Cetacea: Ziphiidae). Proceedings
of the Royal Society, London B. 266: 671-676. Abstract
Hooker, S.K.,
and R.W. Baird. 1999. Observations of Sowerby's beaked whales (Mesoplodon bidens) in The Gully, Nova Scotia. Canadian
Field-Naturalist 113: 273-277. Abstract
Jaquet N., and
H. Whitehead. 1999. Movements, distribution and feeding success of sperm whales
in the Pacific Ocean, over scales of days and tens of kilometers. Aquatic
Mammals 25: 1-13.
Rendell, L.E.,
J. Matthews, A. Gill, J. Gordon and D.W. Macdonald. 1999. Quantitative analysis
of the tonal calls of five odontocete species. Journal of Zoology. 249:
403-410
Smith, S.D.,
and H. Whitehead. 1999. Distribution of dolphins in Galapagos waters. Marine
Mammal Science 15: 550-555. PDF
Whitehead, H.
1999. Testing association patterns of social animals. Animal Behaviour 57: F26-29. PDF
Whitehead, H.
1999. Variation in the visually observable behavior of groups of Galapagos
sperm whales. Marine Mammal Science 15: 1181-1197. PDF
Whitehead, H. 1999. Culture and genetic evolution in whales. Science
284: 2055a PDF
Whitehead, H.,
and S. Dufault. 1999. Techniques for analyzing vertebrate social structure
using identified individuals: review and recommendations. Advances in the
Study of Behavior 28: 33-74. PDF
Publications 1998
Baird, R.W.
1998. An interaction between Pacific white-sided dolphins and a neonatal harbour porpoise. Mammalia 62: 129-134. Abstract
Baird, R.W.,
P.M. Willis, T.J. Guenther, P.J. Wilson and B.N. White. 1998. An intergeneric
hybrid in the family Phocoenidae. Canadian Journal
of Zoology 76: 198-204. Abstract
Bejder, L.,
Fletcher, D. and Brager, S. 1998. A method for testing association patterns of
social mammals. Animal Behaviour 56:719-725. Abstract
Christal, J.,
H. Whitehead and E. Lettevall. 1998. Sperm whale
social units: variation and change. Canadian Journal of Zoology 76:
1431-1440 PDF
Connor,R.C., J. Mann, P.L. Tyack, and H. Whitehead. 1998. Social evolution in
toothed whales. Trends in Ecology and Evolution 13: 228-232. PDF
Dufault, S.,
and H. Whitehead. 1998. Regional and group-level differences in fluke markings
and notches of sperm whales. Journal of Mammalogy 79: 514-520. PDF
Hanson, M.B.
and R.W. Baird. 1998. Dall's porpoise reactions to tagging attempts using a
remotely-deployed suction-cup tag. Marine Technology Society Journal 32(2):
18-23. Abstract
Hooker, S.K.
1998. Extensive scarring observed on female or juvenile male sperm whales off
the Galápagos Islands. Mammalia 62: 134-139. Abstract
Schneider, K.,
R.W. Baird, S. Dawson, I. Visser and S. Childerhouse.
1998. Reactions of bottlenose dolphins to tagging attempts using a
remotely-deployed suction-cup tag. Marine Mammal Science 14: 316-324. Abstract
Whitehead, H.
1998. Cultural selection and genetic diversity in matrilineal whales. Science
282: 1708-1711. PDF
Whitehead, H. 1998.
Male mating strategies: models of roving and residence. Ecological Modelling
111: 297-298. PDF
Whitehead, H. 1998. Cultures of whales and dolphins. WDCS Magazine
20: 4-5.
Whitehead, H.,
M. Dillon, S. Dufault, L. Weilgart and J. Wright.
1998. Non-geographically based population structure of South Pacific sperm
whales: dialects, fluke-markings and genetics. Journal of Animal Ecology
67: 253-262. PDF
Whitehead, H.,
W.D. Bowen, S.K. Hooker, and S. Gowans. 1998. Marine mammals. pp 186 -221 In:
The Gully: a scientific review of its environment and ecosystem. (Eds:
Harrison, W.G. and Fenton, D.G.) Department of Fisheries and Oceans, Ottawa.
Canadian Stock Assessment Secretariat Research Document 98/83.
Willis, P.M.,
and R.W. Baird. 1998. Status of the dwarf sperm whale (Kogia simus) with
special reference to Canada. Canadian Field-Naturalist 112: 114-125. Abstract
Willis, P.M.,
and R.W. Baird. 1998. Sightings and strandings of beaked whales on the west
coast of Canada. Aquatic Mammals 24: 21-25. Abstract
Publications 1997
Baird, R.W.,
and M.B. Hanson. 1997. Status of the northern fur seal, Callorhinus
ursinus, in Canada. Canadian Field-Naturalist 111: 263-269.
Christal, J., and
H. Whitehead. 1997. Aggregations of mature male sperm whales on the Galapagos
Islands breeding ground. Marine Mammal Science 13: 59-69. PDF
Hooker, S.K.,
and R.W. Baird. 1997. A Fea's petrel off Nova Scotia: the first record for Canada.Birders Journal 6: 245-248. Abstract
Paquet, D., C.
Haycock and H. Whitehead. 1997. Numbers and seasonal occurrence of humpback
whales, Megaptera novaeangliae, off Brier Island, Nova Scotia. Canadian
Field-Naturalist 111: 548-552.
Reeves, R. R.,
and H. Whitehead. 1997. Current status of the sperm whale (Physeter
macrocephalus) in Canada. Canadian Field-Naturalist 111: 293-307
Stacey, P.J.,
and R.W. Baird. 1997. Birth of a "resident" killer whale off
Victoria, British Columbia, Canada. Marine Mammal Science 13: 504-508. Abstract
Stacey, P.J.,
D.A. Duffus and R.W. Baird. 1997. A preliminary evaluation of incidental
mortality of small cetaceans in coastal fisheries in British Columbia, Canada. Marine
Mammal Science 13: 321-326.
Weilgart, L., and H. Whitehead. 1997. Group-specific dialects and geographical
variation in coda repertoire in South Pacific sperm whales. Behavioural
Ecology and Sociobiology 40: 277-285.PDF
Whitehead, H. 1997. Analyzing animal social structure. Animal Behaviour 53: 1053-1067. PDF
Whitehead, H.
1997. Sea surface temperature and the abundance of sperm whale calves off the
Galapagos Islands: implications for the effects of global warming. Reports
of the international Whaling Commission 47: 941-944. PDF
Whitehead, H.,
J. Christal, and S. Dufault. 1997. Past and distant whaling and the rapid
decline of sperm whales off the Galapagos Islands. Conservation Biology 11:
1387-1396. PDF
Whitehead, H.,
A. Faucher, S. Gowans, and S. McCarrey. 1997. Status of the northern bottlenose
whale, Hyperoodon ampullatus, in the Gully, Nova Scotia. Canadian
Field-Naturalist 111: 287-292. PDF
Whitehead, H.,
S. Gowans, A. Faucher, and S. McCarrey. 1997. Population analysis of northern
bottlenose whales in the Gully, Nova Scotia. Marine Mammal Science 13:
173-185. PDF
Publications 1996
Baird, R.W.,
and L.M. Dill. 1996. Ecological and social determinants of group size in transient
killer whales. Behavioral Ecology 7: 408-416. Abstract
Jaquet, N., H.
Whitehead and M. Lewis. 1996. Coherence between 19th century sperm
whale distributions and satellite-derived pigments in the tropical Pacific. Marine
Ecology Progress Series 145: 1-10. PDF
Jaquet, N., and
H. Whitehead. 1996. Scale-dependent correlation of sperm whale distribution
with environmental features and productivity in the South Pacific. Marine
Ecology Progress Series135: 1-9. PDF
Richard, K. R.,
H. Whitehead, and J. M. Wright. 1996. Polymorphic microsatellites from sperm
whales and their use in the genetic identification of individuals from
naturally sloughed pieces of skin. Molecular Ecology 5: 313-315.
Richard, K. R.,
M. C. Dillon, H. Whitehead, and J. M. Wright. 1996. Patterns of kinship in
groups of free-living sperm whales (Physeter macrocephalus) revealed by
multiple molecular genetic analyses. Proceedings of the National Academy of
Science (U.S.) 93: 8792-8795. PDF
Weilgart, L., H. Whitehead, and K. Payne. 1996. A colossal convergence. American
Scientist 84: 278-287. PDF
Weller, D.W.,
B. Würsig, H. Whitehead, J.C. Norris, S.K. Lynn, R.W.
Davis, N. Clauss, and P. Brown. 1996. Observations of an interaction between
sperm whales and short-finned pilot whales in the Gulf of Mexico. Marine
Mammal Science 12: 588-594. PDF
Whitehead, H.
1996. Babysitting, dive synchrony, and indications of alloparental care in
sperm whales. Behavioural Ecology and
Sociobiology 38: 237-244. PDF
Whitehead, H. 1996.
Variation in the feeding success of sperm whales: temporal scale, spatial scale
and relationship to migrations. Journal of Animal Ecology 65: 429-438. PDF
Whitehead, H.,
and N. Jaquet. 1996. Are the charts of Maury and Townsend good indicators of
sperm whale distribution and seasonality? Reports of the international
Whaling Commission 46: 643-647.
Publications 1995
Dufault, S.,
and H. Whitehead. 1995. The geographic stock structure of female and immature
sperm whales in the South Pacific. Reports of the international Whaling
Commission 45: 401-405.
Dufault, S.,
and H. Whitehead. 1995. An assessment of changes with time in the marking
patterns used for photo-identification of individual sperm whales, Physeter
macrocephalus. Marine Mammal Science 11: 335-343. PDF
Dufault, S.,
and H. Whitehead. 1995. An encounter with recently wounded sperm whales (Physeter
macrocephalus). Marine Mammal Science 11: 560-563.
Faucher, A., and
H. Whitehead. 1995. Importance of habitat protection for the northern
bottlenose whale in the Gully, Nova Scotia. In Marine protected areas and
sustainable fisheries, ed. N. L. Shackell, and J. H. M. Willison. Wolfville,
Nova Scotia, Canada: Science and Management of Marine Protected Areas
Association.
Gowans, S., and
H. Whitehead. 1995. Distribution and habitat partitioning by small odontocetes
in the Gully, a submarine canyon on the Scotian Shelf. Canadian Journal of
Zoology 73: 1599-1608. PDF
Whitehead, H.
1995. Status of Pacific sperm whale stocks before modern whaling. Reports of
the international Whaling Commission 45: 407-412.
Whitehead, H.
1995. Investigating structure and temporal scale in social organizations using
identified individuals. Behavioral Ecology 6: 199-208. PDF
Whitehead, H.,
and L. Weilgart. 1995. Marine mammal science, the
U.S. Navy and academic freedom. Marine Mammal Science 11: 260-263.
Publications 1994
Dufault, S.,
and H. Whitehead. 1994. Floating marine pollution in 'the Gully' on the
continental slope, Nova Scotia, Canada. Marine Pollution Bulletin 28:
489-493. PDF
Whitehead, H.
1994. Delayed competitive breeding in roving males. Journal of Theoretical
Biology 166: 127-133. PDF
Publications 1993
Baird, R. W.,
P. J. Stacey, and H. Whitehead. 1993. Status of the striped dolphin, Stenella coeruleoalba
in Canada. Canadian Field-Naturalist 107: 455-465.
Dillon, M.C.
and J.M. Wright. 1993. Nucleotide sequence of the D-loop region of the sperm
whale (Physeter macrocephalus) mitochondrial genome. Molecular
Biology and Evolution 10: 296-305.
Dufault, S.,
and H. Whitehead. 1993. Assessing the stock identity of sperm whales in the
eastern equatorial Pacific. Reports of the international Whaling Commission
43: 469-475.
Kahn, B., H.
Whitehead, and M. Dillon. 1993. Indications of density-dependent effects from
comparisons of sperm whale populations. Marine Ecology Progress Series
93: 1-7. PDF
Reeves, R. R.,
E. Mitchell, and H. Whitehead. 1993. Status of the northern bottlenose whale, Hyperoodon
ampullatus. Canadian Field-Naturalist 107: 490-508.
Slooten, E., S.
M. Dawson, and H. Whitehead. 1993. Associations among photographically
identified Hector's dolphins. Canadian Journal of Zoology 71: 2311-2318.
Smith, S. C.,
and H. Whitehead. 1993. Variations in the feeding success and behaviour of Galápagos sperm whales (Physeter
macrocephalus) as they relate to oceanographic conditions. Canadian
Journal of Zoology 71: 1991-1996. PDF
Weilgart, L., and H. Whitehead. 1993. Coda vocalizations in sperm whales (Physeter
macrocephalus) off the Galapagos Islands. Canadian Journal of Zoology
71: 744-752. PDF
Whitehead, H. 1993. The behaviour of mature
male sperm whales on the Galapagos breeding grounds. Canadian Journal of
Zoology 71: 689-699.
PDF
Whitehead, H. and B. Kahn. 1992.
Temporal and geographical variation in the social structure of female sperm
whales. Canadian Journal of Zoology 70: 2145-2149. PDF
Whitehead, H., S. Brennan and D. Grover. 1992.
Distribution and behaviour of male sperm whales on
the Scotian Shelf, Canada. Canadian Journal of Zoology 70: 912-918. PDF
Whitehead, H., S. Waters and T. Lyrholm. 1992. Population structure of female and immature
sperm whales off the Galápagos Islands. Canadian Journal of Fisheries and
Aquatic Sciences 49: 78-84.
Whitehead, H. and S. Walde. 1992. Habitat
dimensionality and mean search distance of top predators: implications for
ecosystem structure. Theoretical Population Biology 42: 1-9. PDF
Amos, W., H. Whitehead, M.J. Ferrari, D.A.
Glockner-Ferrari, R. Payne and J. Gordon. 1992. Restrictable
DNA from sloughed cetacean skin; its potential for use in population analysis. Marine
Mammal Science 8: 275-283.
Hope, P.L. and H. Whitehead. 1991. Sperm whales
off the Galápagos Islands during the period 1830-1850 and comparisons with
modern studies. Reports of the International Whaling Commission 41:
273-283.
Whitehead, H. and L.S. Weilgart.
1991. Patterns of visually observable behaviour and
vocalizations in groups of female sperm whales. Behaviour
118: 275-296. PDF
Whitehead, H., S. Waters and T. Lyrholm. 1991. Social organization in female sperm whales
and their offspring: constant companions and casual acquaintances. Behavioral
Ecology and Sociobiology 29: 385-389. PDF
Whitehead, H. and P.L. Hope. 1991. Sperm
whalers off the Galápagos Islands and in the northwest Pacific, 1830-1850:
ideal free whalers? Sociobiology and Ethology 12: 146-161.
Whitehead, H. 1990. Rules for roving males. Journal
of Theoretical Biology 145: 355-368.
Whitehead, H. and L. Weilgart.
1990. Click rates from sperm whales. Journal of the Acoustical Society of
America 87: 1798-1806.
Whitehead, H. 1990. Mark-recapture estimates
with emigration and re-immigration. Biometrics 46: 473-479.
Weilgart, L.S. and H. Whitehead. 1990.
Vocalizations of the North Atlantic pilot whale (Globicephala melas) as
related to behavioral contexts. Behavioral Ecology and Sociobiology 26:
399-402.
Whitehead, H., J. Gordon, E.A. Mathews, and
K.R. Richard. 1990. Obtaining skin samples from living sperm whales. Marine
Mammal Science 6: 316-326. PDF
Whitehead, H. 1990. Assessing sperm whale
populations using natural markings: a progress report. Reports of the
International Whaling Commission (Special Issue 12): 377-382. PDF
Waters, S. and H. Whitehead. 1990. Aerial behaviour in sperm whales, Physeter macrocephalus. Canadian
Journal of Zoology 68: 2076-2082.
Whitehead, H. and S. Waters. 1990. Social
organization and population structure of sperm whales off the Galápagos
Islands, Ecuador (1985 and 1987). Reports of the International Whaling
Commission (Special Issue 12):
249-257. PDF
Carlson, C., C. Mayo and H. Whitehead. 1990.
Changes in the ventral fluke patterns of individually identified humpback
whales. Reports of the International Whaling Commission (Special Issue 12):
105-111.
Whitehead, H. 1990. Computer-assisted
individual identification of sperm whale flukes. Reports of the International
Whaling Commission (Special Issue 12): 71-77.
Waters, S. and H. Whitehead. 1990. Population
and growth parameters of Galápagos sperm whales estimated from length
distributions. Reports of the International Whaling Commission 40: 225-235.
Whitehead, H., L. Weilgart
and S. Waters. 1989. Seasonality of sperm whales off the Galápagos Islands,
Ecuador. Reports of the International Whaling Commission 39: 207-210.
Whitehead, H. 1989. Foraging formations of
Galápagos sperm whales. Canadian Journal of Zoology 67:
2131-2139. PDF
Papastavrou, V., S.C. Smith and H.
Whitehead. 1989. Diving
behaviour of the sperm whale, Physeter
macrocephalus, off the Galápagos Islands. Canadian Journal of Zoology
67: 839-846. PDF
Arnbom, T. and H. Whitehead. 1989.
Observations on the composition and behaviour of
groups of female sperm whales near the Galápagos Islands. Canadian Journal of Zoology
67: 1-7. PDF
Whitehead, H., V. Papastavrou and S. Smith.
1989. Feeding success of sperm whales and sea-surface temperatures off the
Galápagos Islands. Marine Ecology
Progress Series 53: 201-3.
Whitehead, H. 1989. Voyage to the whales. Stoddart, Toronto
Katona, S.K. and H. Whitehead. 1988. Are
cetaceans ecologically important? Oceanography
and Marine Biology Annual Review 26: 553-568. PDF
Mullins, J., H. Whitehead and L.S. Weilgart. 1988. Behaviour and
vocalizations of two single sperm whales, Physeter macrocephalus,
off Nova Scotia. Canadian Journal of Fisheries and Aquatic Sciences 45:
1736-43. PDF
Weilgart, L.S. and H. Whitehead. 1988.
Distinctive vocalizations from mature male sperm whales (Physeter
macrocephalus). Canadian Journal of Zoology 66:
1931-7. PDF
Whitehead, H. and C. Carlson. 1988. Social behaviour of feeding finback whales off Newfoundland:
comparisons with the sympatric humpback whale. Canadian Journal of Zoology
66: 217-221. PDF
Alling, A. and H. Whitehead. 1987. A
preliminary study of the status of the whitebeaked
dolphin and other small cetaceans off the coast of Labrador. Canadian
Field-Naturalist 101: 131-135. PDF
Arnbom, T., V. Papastavrou, L.S. Weilgart and H. Whitehead. 1987. Sperm whales react to an
attack by killer whales. Journal of
Mammalogy 68: 450-453. PDF
Whitehead, H. 1987. Updated status of the
humpback whale, Megaptera novaeangliae, in Canada. Canadian
Field-Naturalist 101: 284-294.
Whitehead, H. and T. Arnbom.
1987. Social organization of sperm whales off the Galápagos Islands,
February-April 1985. Canadian Journal of Zoology 65(4):
913-919. PDF
Rice, M., C. Carlson, K. Chu, W. Dolphin and H.
Whitehead. 1987. Are humpback whale population estimates being biased by sexual
differences in fluking behavior? Reports
of the International Whaling Commission 37: 333-335.
Whitehead, H. 1987. Social organization of
sperm whales off the Galápagos: implications for management and conservation. Reports
of the International Whaling Commission 37: 195-199.
Whitehead, H. 1987. Sperm whale behavior on the
Galápagos Grounds. Oceanus 30: 49-53.
Whitehead, H., R. Payne and M. Payne. 1986.
Population estimate for the right whales off Peninsula Valdes, Argentina,
1971-1976. Reports of the International Whaling Commission (Special Issue 10): 169-171.
Whitehead, H. and J. Gordon. 1986. Methods of
obtaining data for assessing and modelling sperm whale populations which do not
depend on catches. Reports of the International
Whaling Commission (Special Issue 8): 149-166.
Weilgart, L.S. and H. Whitehead. 1986.
Observations of a sperm whale (Physeter
catodon) birth. Journal of Mammalogy
67: 399-401. PDF
Whitehead, H. and C. Glass. 1985. The
significance of the Southeast Shoal of the Grand Bank to humpback whales and
other cetacean species. Canadian Journal of Zoology 63:
2617-2625. PDF
Whitehead, H. and C. Glass. 1985. Orcas (killer
whales) attack humpback whales. Journal of Mammalogy 66: 183-185.
Whitehead, H. 1985. Why whales leap. Scientific
American 252: 84-93. PDF
Whitehead, H. and J.E. Carscadden. 1985.
Predicting inshore whale abundance - whales and capelin off the Newfoundland
coast. Canadian Journal of Fisheries and Aquatic Sciences 42:
976-981. PDF
Whitehead, H. 1985. Humpback whale songs from
the northern Indian Ocean. Investigations
on Cetacea (Berne, Switzerland) 17: 157-162. PDF
Whitehead, H. 1985. Humpback whale breaching. Investigations on Cetacea (Berne,
Switzerland) 17: 117-155.
PDF
Whitehead, H. 1985. Benign research on sperm
whales in the Indian Ocean Sanctuary. Journal of the American Cetacean Society
19: 3-7.
Whitehead, H. 1985. Studying sperm whales on
the Galápagos Grounds. Noticias de Galápagos 42: 18-21.
Whitehead, H., K. Chu, J. Perkins, P. Bryant
and G. Nichols. 1983. Population size, stock identity, and distribution of the
humpback whales off West Greenland - Summer 1981. Reports of the International
Whaling Commission 33: 497-501.
Whitehead, H. 1983. Structure and stability of
humpback whale groups off Newfoundland. Canadian Journal of Zoology 61:
1391-97. PDF
Tyack, P. and H. Whitehead. 1983. Male
competition in large groups of wintering humpback whales. Behaviour
83: 132-154.
Whitehead, H. 1983. Baleen whales off Sri
Lanka. Loris 16: 176-181. PDF
Whitehead, H. 1982. Sperm whales off
Sri Lanka. Loris 16: 23-24. PDF
Whitehead, H., K. Chu, P. Harcourt and A.
Alling. 1982. The humpback whales off West Greenland: summer 1981. Final Report
to Marine Mammal Commission. N.T.I.S. Refereed Publication no.
PB82-243924.
Whitehead, H. 1982. Populations of humpback
whales in the northwest Atlantic. Reports of the International Whaling
Commission 32: 345-353.
Whitehead, H., P. Harcourt and R. Silver. 1982.
The migration of the humpback whales along the northeast coast of Newfoundland.
Canadian
Journal of Zoology 60: 2173-9.
PDF
Whitehead, H. and M.J. Moore. 1982.
Distribution and movements of West Indian humpback whales in winter. Canadian
Journal of Zoology 60: 2203-11.
PDF
Whitehead, H., and R. Payne. 1981. New
techniques for measuring whales from the air. Final Report to Marine Mammal
Commission. N.T.I.S. Refereed Publication
no. PB81-161143. 36 pp.
Whitehead, H. 1981. The humpback whales in the
Northwest Atlantic: behaviour, ecology and
populations. In: Proceedings of a Workshop on Humpback Whales of the western North
Atlantic, Boston, Nov. 1980. Final
Report to N.M.F.S. Appendix II-B: 35-58.
Whitehead, H. and R. Payne. 1981. New
techniques for assessing populations of right whales without killing them. In: Mammals of the Sea, Vol. 3. FAO Fish.
Ser. 5, Rome, Italy: 189-209.
Katona, S.K., and H.P. Whitehead. 1981.
Identifying humpback whales using their natural markings. Polar Record 20: 439-444.
Whitehead, H., P. Harcourt, K. Ingham, and H.
Clark. 1980. Migration of the baleen whales past the Bay de Verde Peninsula,
Newfoundland. Canadian Journal of Zoology 58: 687-692.
Katona, S., B. Baxter, O. Brazier, S. Kraus, J.
Perkins and H. Whitehead. 1979. Identification of humpback whales by fluke
photographs. In: Behavior of Marine
Animals, Vol. 3. Edited by H.E. Winn
and B.L. Olla. Plenum Press, New York:
33-44.
Perkins, J. and H. Whitehead. 1977. Observations
on three species of baleen whales off northern Newfoundland and adjacent
waters. Journal of the Fisheries Research Board of Canada 34:
1436-1440. PDF
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